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Cajaraville (1995) The effect of sublethal lead exposure on the ultrastructure and on the distribution of acid phosphatase activity in chloragocytes of earthworms (Annelida, Oligochaeta) Histochemical Journal (London), 27, 965 973 Cancio, I, A Orbea, A V lkl, HD Fahimi & MP Cajaraville (1998) Induction of peroxisomal oxidases in mussels: comparison of effects of lubricant oil and benzo[a]pyrene with two typical peroxisome proliferators on peroxisome structure and function in Mytilus galloprovincialis Toxicology and Applied Pharmacology, 149, 64 72 Cancio, I, A Ibabe & MP Cajaraville (1999) Seasonal variation of peroxisomal enzyme activities and peroxisomal structure in mussels Mytilus galloprovincialis and its relationship with the lipid content Comparative Biochemistry and Physiology, 123, 135 144 Carginale, V, R Scudiero, C Capasso, A Capasso, P Kille, G di Prisco & E.

3 Possible causes for M74 The symptoms of the disease occur in newly hatched yolk-sac fry during the phase when the yolk sac is resorbed before the fry start to feed The fry initially show hyperactivity followed by loss of co-ordination, hyperpigmentation, precipitates in yolk sac apathy and exophthalmia Death occurs usually a few days after the rst external symptoms The symptoms are female-speci c; that is, the M74 affects offspring of certain females with total mortality As indicated above, it is by now well-established that the immediate cause for M74 in a sh is a lack of thiamine (vitamin B1) and female salmon have been routinely treated with thiamine since 1994 Treatment with thiamine has been shown to protect both adult and fry from M74 It is by no means entirely clear why Baltic salmon are de cient in thiamine in some years and not in others.

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And this conditional loss rate can be used to attribute capital to asset A : Conditional Attribution to A = (Conditional Loss Rate of A ) LGD(A) To implement this, we need to de ne that threshold. Suppose that, for the determination of economic capital for this portfolio, we are using a

Parisi (1998) Cadmium-induced differential accumulation of metallothionein isoforms in the Antarctic ice sh, which exhibits no basal metallothionein protein but high endogenous mRNA levels Biochemical Journal, 332, 475 481 Carpen , E & M Vasak (1989) Hepatic metallothioneins from gold sh (Crassius auratus L) Comparative Biochemistry and Physiology, 92B, 463 468 Carrano, A, VLH Thompson, PA Lindl & JL Minkler (1978) Sister chromatid exchange as an indicator of mutagenesis Nature, 271, 551 553 Casciano, DA (1991) Introduction: historical perspectives of genetic toxicology In: (eds Li, AP & RH He ich) Genetic Toxicology CRC Press, Boca Raton, Florida, pp 1 12 Cattley, RC, JG Conway & JA Popp (1987) Association of persistent peroxisome proliferation and oxidative injury with hepatocarcinogenicity in female F-34 rats fed Di(2-ethylhexyl)phthalate for 2 years Cancer letters, 38, 15 22 Chang, L-Y, J.

Possible causal factors include food availability and selection, exposure to xenobiotics, infectious agents, endogenous levels of antioxidants, activity of antioxidant enzymes and low genetic diversity Each of the factors mentioned may interact and could in its turn be affected by environmental in uences such as eutrophication, acid precipitation and over shing of salmon or other Baltic species In both EMS and M74 the diet appears to be an important parameter Thiamine cannot be synthesised by higher organisms and must therefore be taken in through diet (or water) or produced by endosymbiontic microorganisms (Cooray et al, 1999) Shifts in the diet of salmonids from crustaceans and sh species low in thiaminase to thiaminase-rich sh species appear to be important factors in the development of the diseases (Fitzsimons et al, 1999; Karlsson et al, 1999).

W Slot & HJ Geuze (1988) Molecular immunocytochemistry of the CuZn superoxide dismutase in rat hepatocytes Journal of Cell Biology, 107, 2169 2179 Cheng, R, BL Ford, PE Oneal, CZ Mathews, CS Bradford, T Thongtan, DW Barnes, JD Hendricks & GS Bailey (1997) Zebra sh (Danio rerio) p53 tumor suppressor gene: cDNA sequence and expression during embryogenesis Molecular Marine Biology and Biotechnology, 6, 88 97..

In the Baltic, different diets could also explain why some salmon populations are not affected..

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